The One, also known as the Source and the sentient core of the universe, is an extradimensional entity from before time, in whom Order and Chaos exist in perfect harmony. He is the grand creator of the Tranformers Multiverse. It created Unicron, then cut him in half to make Primus when Unicron began consuming everything and had Primus destroy him before he could do any further harm. It then left the multiverse in charge of Primus and it proceeded to continue its endless travel through reality, spreading life and balance.
The Vok claim to be guardians of The One and followers of its will. They also seem to think that either it created the multiverse, or it is the multiverse.